GALIUM TRICORNUTUM

Family and Genus:- See- RUBIACEAE/Sect. KOLGYDA

Common Names:- Corn cleavers.

Homotypic Synonyms:-  Galium pisiferum, Galium tricorne

Meaning:- Galium (Gr) Milk.
                  Tricornutum (L) With three horns.

General description:- Annual.

Stems:-
1) 10-80(-100) cm, scrambling, stout, retrorsely aculeolate, scabrous, without
    straight hairs below and above the nodes.

Leaves:-
1) 10-40 x 2·5-8 mm, in whorls of 6-8, narrowly oblanceolate, long-awned, glabrous,
    above, the margin and midrib retrorsely scabrid); margins revolute.

Flower:-
1) Inflorescence, long;
    a) partial inflorescences, (1-)3- to 5(-7)-flowered, scarcely longer than the leaves;
2) Lower peduncles, (5-)8-15(-20) mm.
3) Pedicels, 1-8 mm, curved distinctly inwards and downwards after flowering.
4) Flowers, hermaphrodite, the lateral often male.
5) Corolla, 1-1·7 mm diam. glabrous, white;
    a) lobes, long, acute.

Fruit:-
1) Mericarps, 3-4(-5) mm, densely covered with small, acute papillae.

Key features:-
1) Leaf-margin, retrorsely aculeolate; leaves glabrous above.
2) Fruits, more than 2 mm. verrucose.
3) Peduncles and pedicels, convergent and deflexed after anthesis,

Click here for a glossary of terms used

Habitat:- A weed of traditional argiculture, with Ranuculus arvensis, Biflora
testiculata, Agrostema githargo, etc. occasionally in gorges and open woodland. 0-
1200 m.

Distribution:- Throughout mainland Greece. - Mediterranean region and SW Asia.
Somewhat sparsely scattered across Crete not common.

Flowering time:- Late Mar to early June.

Photos by:- Steve Lenton
SPECIES DESCRIPTION