POLYCARPON TETRAPHYLLUM

Family:- CARYOPHYLLACEAE

Common Names:- Four-leaved allseed

Synonyms:- Holosteum tetraphyllum, Polycarpaea tetraphylla, Mollugo
tetraphylla

Meaning:- Polycarpon (Gr) Many-fruited, a name used by the Greek physician
Hippocrates.
                  Tetraphyllum (L) Four leaved, with leaves in fours.
                
General description:- Glabrous or somewhat papillose annual or biennial.

Stems:-
1) 4-12 cm, usually much branched.

Leaves:-
1) Opposite or in whorls of 4, 4-15 mm, elliptical to broadly oblanceolate.
2) Stipules and bracts 2-3 mm, triangular-ovate, acuminate, scarious.

Flowers:-
1) Inflorescence lax, spreading, in branched clusters, white, tiny, 2-3 mm.
2) Petals, emarginate, narrow, shorter than the sepals.
3) Sepals, c. 2 mm, ovate, keeled, with whitish margins.
4) Stamens, (1-)3-5.

Fruit:-
1) Capsule, dehiscing, with 3 valves almost to the base; valves twisting spirally into
    tubes.
2) Seeds c. 0.5 mm, several, minutely tuberculate.

Key features:-
1) Annual to perennial, without a woody stock.
2) Stipules and bracts ± conspicuous and silvery.
3) Petals usually emarginate.

Habitat:- Dry, sandy and gravelly places, coastal habitats, streambeds, roadsides,
fallow fields and wasteground. 0-800(-1700)m.

Distribution:- Widespread throughout the Mediterranean region SW Europe and
SW Asia. Widespread on Crete.

Flowering time:- Mainly Apr-June.

Photos by:- Steve Lenton
SPECIES DESCRIPTION
 
GLOSSARY OF TERMS USED

Acuminate:- Gradually narrowing to a point.

Capsule:- Dry fruit that opens when ripe. splitting from the apex to the base into
separate segments known as valves.

Bract:-  An organ, often small and scale-like, but sometimes leaf-­like, located
where the flower-stalk joins the stem.

Dehiscent - Dehiscing:- Splitting open to release the seeds.

Elliptic - Elliptical:- Forming an ellipse, widest in the middle and pointed at both
ends.
Emarginate:- Distinctly notched at the apex.

Glabrous:- Without hairs, hairless.

Inflorescence:- The flowering branch or branches, flowers and bracts above the
uppermost leaves on a stem. Inflorescences are very variable from one species to
another.

Keel - Keeled:- A prominent longitudinal ridge like the keel of a boat.

Oblanceolate:- Inversely lanceolate, broadest towards the apex and tapering to
the stalk.
Ovate:- Broad and rounded at the base and tapering toward the end.

Papillose:- Covered with papillae. covered  with small nipple-like projections.
Petal:- The inner perianth segments when they clearly differ from the outer.

Scarious:- Thin and dry, paper-like, membranous not green.
Sepal:- A member of the outer perianth whorl in most flowers. The sepals
collectively make up the calyx.
Stamen:- Pollen-producing reproductive organ, typically consisting of a stalk called
the filament and an anther.
Stipule:- An outgrowth typically borne on both sides (sometimes on just one side)
of the base of a leafstalk (the petiole).

Triangular-ovate:- Egg-shaped, with the broader end at the base.
Tuberculate:- With small, wart-like projections.
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